NEW YORK — Jacksonville Jaguars defensive tackle Malik Jackson was fined $24,309 by the NFL on Friday after being penalized twice for unsportsmanlike conduct during one drive last weekend.

Jackson was called for roughing the passer on Oakland's Derek Carr, then he protested the call and was penalized for using abusive language toward an official. Four plays later, Jackson was again called for unsportsmanlike conduct after a field goal and then ejected. Under new NFL rules, players are tossed if they are penalized twice for unsportsmanlike conduct in the same game.

Wide receiver Marqise Lee, Jackson's teammate, was docked $12,154 on Friday after he received an unsportsmanlike conduct penalty for using a racial slur in the Jaguars' 33-16 loss.

Miami receiver Jarvis Landry was fined $24,309 for an illegal crackback block, as reported previously by The Associated Press , that sent Buffalo's Aaron Williams to the hospital with a head and neck injury.

The Dolphins' Byron Maxwell was hit with a $9,115 fine for a taunting penalty. The Bills' Richie Incognito was also docked the same amount for throwing the helmet of Miami defensive end Andre Branch during a fourth-quarter scuffle.

Baltimore's Za'Darius Smith was fined $18,231 for roughing the passer on the New York Jets' Ryan Fitzpatrick.

Giants defensive end Owa Odighizuwa has to pay $12,154 after he pretended to take a picture in the end zone of safety Landon Collins, who returned an interception for a touchdown against Los Angeles in London last Sunday. Odighizuwa was penalized for unsportsmanlike conduct on the play.

Minnesota guard Alex Boone was docked $12,154 for unsportsmanlike conduct after yelling at the refs during the Vikings' loss to Philadelphia.

San Francisco defensive end DeForest Buckner was fined $9,115 for unnecessary roughness in the 49ers' loss to Tampa Bay.

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AP NFL website: www.pro32.ap.org and AP NFL Twitter feed: www.twitter.com/AP_NFL